S24.003] Gene Transfer Clinical Trial for Limb Girdle Muscular Dystrophy Type 2D (Alpha-Sarcoglycan Deficiency)
Jerry R. Mendell, Hansell Stedman, Steven A. Moore, Valérie Allamand, Kevin P. Campbell, Cheryl Wall, Cathy Stolle, Isabelle Periquet, Vera Novak, James W. Wilson Columbus , OH; Philadelphia, PA; Iowa City , IA

OBJECTIVE: Determine the safety of gene transfer in patients with limb girdle muscular dystrophy (LGMD) type 2D (alpha-sarcoglycan deficiency)
BACKGROUND: Current treatment opportunities for muscular dystrophy are limited. LGMD caused by sarcoglycan deficiencies are particularly suitable for viral-mediated gene transfer to skeletal muscle using adeno-associated virus (AAV). This replication defective, nonpathogenic vector combines high-efficiency with low immunogenicity and can transfer the entire cDNA of the sarcoglycan genes. AAV has been used to successfully transfer the sarcoglycan and dystrophin coding sequences in animals.
DESIGN/METHODS: Two subjects participated in this trial, a 37-year-old man (patient 1) and a 14-year-old boy (patient 2). Both had missense mutations of the alpha-sarcoglycan (a-SG) gene. Expression of a-SG in quadriceps muscle biopsies prior to gene transfer was greatly reduced in patient 1 and absent in patient 2. The full length a-SG cDNA was cloned into an AAV-2 vector plasmid containing a cytomegalovirus (CMV) promoter to generate the vector construct, AAV-CMV-a-SG. This construct was shown to express a-SG in a mouse model of LGMD 2D. The extensor digitorum brevis (EDB) was chosen as the site of gene transfer and following a double-blind, randomized protocol, one side received vector and the other side saline. Forty-three days after gene transfer the muscles were biopsied for analysis.
RESULTS: Both patients tolerated the injections into the EDB without adverse effects. Neither developed signs of local or systemic reaction at any time. No differences could be discerned in histopathology comparing the right and left EDBs of either patient. Dystrophic changes were observed on both sides. Although macrophages infiltrated muscle fibers undergoing necrosis, no endomysial or perimysial inflammatory cell infiltrates were observed. Results of gene expression will be shown at the time of presentation.
CONCLUSIONS: Several important lessons were learned from this first gene transfer to skeletal muscle in muscular dystrophy. The patients experienced no adverse reactions and there were no signs of inflammation detected in the muscle biopsies post gene transfer. This sets the stage for future studies using AAV in clinical gene transfer trials in musuclar dystrophy. The EDB posed problems in interpretation related to retained sarcoglycan expression in these patients with missense mutations of the a- SG gene. In future gene transfer trials limb muscles may improve opportunities for assessing efficacy both in regard to gene expression and for measurement of strength using quantitative methods.
Supported By: Muscular Dystrophy Association

P01.069] Peak Expiratory Flow (PEF) Is a Useful Measure of Muscle Strength in Young Boys with Duchenne Muscular Dystrophy (DMD)
Richard I. Webster, Alison Boynton, Richard L. Henry, Heather M. Johnston Randwick, New South Wales, Australia

OBJECTIVE: To evaluate the use of Peak Expiratory Flow (PEF) as a measure of muscle strength in young boys with Duchenne Muscular Dystrophy (DMD) and to compare the use of PEF with other measures of respiratory function.
BACKGROUND: There is a need to measure the effect of treatments that potentially increase muscle strength in DMD. Assessment of muscle strength in young boys with DMD is often hampered by problems with attention and concentration. Vital Capacity (VC) is the most commonly used measure of respiratory function but young boys find it a difficult manoeuvre. PEF is easily performed, correlates with other measures of respiratory muscle strength and so potentially has some advantages as a measure of muscle strength.
DESIGN/METHODS: 17 boys with DMD (5-10 years of age) were seen at 4 monthly intervals over one year. Each assessment involved respiratory function testing (PEF, VC, one second Forced Expiratory Volume [FEV1]), timed functional testing (walking 9 metres, climbing 4 stairs and arising from supine) and manual muscle strength testing (MMST). A single investigator performed MMST and timed functional testing. A separate investigator performed respiratory function testing. Both investigators were blinded to the results of the other's assessment. During the study 11/17 boys were treated with prednisolone, which increases muscle strength in DMD.
RESULTS: At baseline, mean PEF (69% of predicted) and VC (76% of predicted) were abnormal. PEF and VC manoeuvres were assessed as being adequately performed in 80% and 65% of assessments respectively. Only 22% of flow-volume loops were considered to accurately estimate VC. At baseline, there was a significant correlation between PEF (% predicted) and strength assessed by MMST (P=0.002) and between PEF (% predicted) and the time to walk 9 metres (P=0.01). At baseline, neither VC (% predicted) nor FEV1 (% predicted) showed a statistically significant correlation with MMST or timed functional testing. At the time of the final assessment, change from baseline was calculated for each boy. There was a significant correlation between the change in PEF (% predicted) and the change in: MMST (P=0.03), time to climb 4 stairs (P=0.001), time to walk 9 metres (P=0.001) and time to arise from supine (P=0.002). Neither change in VC (% predicted) nor FEV1 (% predicted) showed a statistically significant correlation with these parameters. Boys treated with prednisolone did better than those not treated, in all parameters. PEF, time to walk 9 metres and time to climb 4 stairs showed the only statistically significant improvement. Mean PEF (% predicted) showed a 20% improvement from baseline in the treated boys and a 2% improvement in the untreated boys (P=0.007).
CONCLUSIONS: PEF is abnormal early in the course of DMD, correlates well with other measures of muscle strength and is sensitive to changes in muscle strength induced by prednisolone. It is more easily performed than spirometry and hence has some advantages as a means of monitoring muscle strength in young boys with DMD.
Supported By: The Muscular Dystrophy Association of New South Wales
 

P04.147] Integrin alfa7beta1 in Muscular Dystrophy/Myopathy of Unknown Etiology
Elena Pegoraro, Paola Prandini, Marina Fanin, Guido Tarone, Eva Engvall, Corrado Angelini Padova, Italy; Torino, Italy; La Jolla, CA

OBJECTIVE: To investigate the role of integrin alfa7 in muscle pathology we studied a large series of muscle biopsy from patients affected with unclassified muscular dystrophy/myopathy.
BACKGROUND: Integrins are transmembrane heterodimer glycoproteins resulting from the assembly of two different subunits, a and b, and have a prominent role in myogenesis, differentiation, cell migration and cell-cell interactions. The alfa7 subunit is mainly expressed in skeletal and cardiac muscle and it has been previously shown to be primarily mutated in three patients affected with congenital myopathy with variable clinical phenotype.
DESIGN/METHODS: We used a candidate gene and protein approach in a large cohort of muscular dystrophy/myopathy patients. Antibody against the intracellular domain of the integrin alfa7 was used to immunostain muscle biopsy from 210 patients with muscular dystrophy/myopathy of unknown etiology. Patients whose muscle biopsy specimen showed integrin a7 deficiency on immunostaining were studied for integrin alfa7 gene mutations with reverse transcription of muscle RNA analysis involving single strand conformational polymorphism and sequencing.
RESULTS: Levels of alfa7 integrin were found to be decreased on immunostaining of muscle biopsy specimen from 35 of the 210 patients (~17%). In six of these patients no integrin a7 was detected. Screening for alfa7 mutation in 30 of the 35 patients detected only one integrin alfa7 missense mutation (the mutation on the second allele was not found) in a patient presenting with a congenital muscular dystrophy (CMD)-like phenotype (floppy infant with multiple joint contractures and abnormal white matter signal at brain CT scan). No integrin alfa7 gene mutations were identified in all the other patients showing integrin alfa7 deficiency. Moreover, we identified a novel integrin alfa7 isoform presenting 71 bp deletion. This isoform results from a partial deletion of exon 21 due to the use of a cryptic splice site generated by a G to A missense mutation at nucleotide position 2644 in integrin alfa7 cDNA. This spliced isoform is present in about 12% of the chromosomes studied.
CONCLUSIONS: Our results suggest that secondary integrin alfa7 deficiency is rather common in muscular dystrophy/myopathy of unknown etiology. This is not surprising considering the central role played by alfa7beta1 integrin in anchoring the intracellular cytosketon via actin, to the extracellular matrix, via laminin, and the potential for signal transduction possibly mediated by extracellular matrix ,soluble growth factors, and/or associated transmembrane molecules. Thus, several distinct primary defects may converge into an integrin common pathway system determining underexpression of integrin as a modulatory phenomenon.
Supported By: Telethon grant # 1114
 

S24.004] Novel Sarcoglycan Mutations Widen the Clinical Spectrum of Limb-Girdle Muscular Dystrophy 2C,2D,2E,2F
Corrado I. Angelini, Chiara Boito, Marina Fanin, Gabriele Siciliano, Elena Pegoraro Padova, Italy; Pisa, Italy

OBJECTIVE: To identify primary sarcoglycan mutations in a group of 221 autosomal recessive limb-girdle muscular dystrophy (LGMD) and myopathy patients.
BACKGROUND: A new clinical molecular classification of Limb-girdle muscular dystrophy has been proposed: the defect of four transmember glycoproteins alpha, beta, gamma, delta sarcoglycan results in LGMD type 2D, 2D, 2E, 2C and 2F respectively. All sarcoglycan defects cause the loss of SG (sarcoglycan) complex and are due to very heterogeneous set of mutations that cause different molecular consequences. The loss of SG complex leads to membrane instability in skeletal and cardiac muscle. A possible mechanism has been identified since SG complex deficiency leads to the unmasking of metallo-proteases cleavage site on beta-dystroglycan, causing the disruption of the link between the basement of membrane and sarcolemma.
DESIGN/METHODS: Muscle biopsy of 221 LGMD/myopathy patients were analized by immunohistochemistry and immunoblot to study the expression of four subunits of SG complex (alpha,beta, gamma, delta SG). Abnormal SG expression was found in 24 cases. RNA was extracted from biopsies and was studied by RT-PCR and SSCP analysis, we identified the mutations by cDNA direct sequencing.
RESULTS: We found 7 patients with novel frameshift and missense mutations in alpha-SG: their clinical phenotype ranged from hyperCKemia to mild myopathy or severe LGMD. One patient showed an homozygous out of frame duplication of 32 base pair in beta-sarcoglycan gene and presented a severe dilated cardiomyopathy, 3 patients had mutation in gamma-sarcoglycan and presented severe muscular dystrophy. We found a novel delta-SG mutation in a female with a limb-girdle phenotype.
CONCLUSIONS: Our data combined with our previous screening study suggest that in our patients population sarcoglycanopathy account for about 15% of LGMD cases. The most frequent mutations were in alpha-SG which result in a spectrum of clinically heterogeneous phenotypes ranging from a rapid progressive to a late onset slow course. We describe a new further delta-sarcoglycan mutation causing a mild LGMD phenotype. In our study the more severe clinical phenotypes were in LGMD 2E and 2D. Dilated cardiomyopathy was prominent in beta-SG patient and widens the clinical phenotype.This might be explained by the involvement of vascular smooth muscle, were beta-sarcoglycan is present, leading to coronary disfunction and myocardial necrosis.
Supported By: Telethon project n.1321-1114,C.54.

 P01.062] A New, Highly Efficient Fully "Gutted" Adenoviral Recombinant for Dystrophin Gene Transfer to Newborn and Adult Muscle
Renald Gilbert, An-Bang Liu, Basil J. Petrof, Josephine Nalbantoglu, George Karpati Montreal , QC, Canada; Hualien, Taiwan

OBJECTIVE: To construct a fully gutted adenovirus (Ad), also known as helper-dependent Ad (HDAd), for efficient and long-term expression of dystrophin in muscle.
BACKGROUND: HDAd hold great promises as a gene transfer vector for the treatment of genetic diseases such a Duchenne muscular dystrophy. They do not encode any viral genes and are consequently less immunogenic than E1-deleted Ad (first generation Ad). Moreover, HDAd have an increased transport capacity and can carry up to 35 kb of foreign DNA. In addition to the vector, the intensity and duration of the transferred gene expression is greatly influenced by the promoter. Therefore we investigated a hybrid promoter (cytomegalovirus enhancer/b-actin promoter), known as CB, whose efficiency is expected to be superior to any previous promoters employed in the context of a HDAd. To further increase the amount of transgene protein produced, two tandem dystrophin expression cassettes regulated by the CB promoter were inserted into the backbone of the HDAd.
DESIGN/METHODS: A HdAd without any viral genes and encoding two human full-length dystrophin cDNAs (12 kb each) regulated by the CB promoter was constructed and called HDCBDysDys. This vector was amplified by several passages on 293cre cells and purified by centrifugation on CsCl gradients. The tibialis anterior (TA) muscle of neonatal and young adult dystrophin-deficient mdx mice was injected with 5 ml or 30 ml respectively of HDCBDysDys at a titer of 1.0 X 10
12 virus particles/ml. At 10 and 30 days (adults) or at 10, 30 and 90 days post-injection (neonatals), the injected muscles were analyzed by immunohistochemistry for dystrophin expression. The sera of some animals were also analyzed by western blot for the presence of antibodies against dystrophin.
RESULTS: At 10 days post-injection, the average number of dystrophin positive fibers/muscle was 582 (34% of total) and 104 (6.1% of total) for neonatal and adult mdx mice respectively. In addition, many transduced fibers demonstrated an intense dystrophin positive cytoplasmic staining, a sign of marked dystrophin over-expression. This transduction level was significantly higher than the one observed after gene transfer with a HDAd encoding a single dystrophin expression cassette regulated by the cytomegalovirus promoter (Gilbert et al., Hum. Gene Ther. 12:1741). However, significant reduction in the number of dystrophin positive fibers was observed after 30 and 90 days in the two animal groups. Antibodies against the human dystrophin transgene were demonstrated in the sera of some animals, indicating the transgene protein was immunogenic.
CONCLUSIONS: Very high but transient dystrophin expression was observed after gene transfer with HDCBDysDys. It remains unclear if the progressive reduction of the transgene expression was due to the antigenicity of dystrophin or to other factors unrelated to the immune response. These questions are being investigated by repeating the previous experiments in immunodeficient mice and by generating a HDAd encoding the murine dystrophin.
Supported By: Muscular dystrophy association of USA and Canada

P01.065] Efficient and Long-Term Unabated Gene Expression in Immunodeficient Muscle Using a Partially Gutted Adenovirus Vector
Renald Gilbert, Jatinderpal R. Deol, An-Bang Liu, Joon-Shik Moon, Basil J. Petrof, Josephine Nalbantoglu, George Karpati Montreal , QC, Canada; Hualien, Taiwan

OBJECTIVE: To test if retention of two adenoviral genes i.e. E1B and E4 in the backbone of an otherwise gutted adenovirus (Ad) will produce and efficient and prolonged transgene expression.
BACKGROUND: Helper-dependent Ad (HDAd), also known as fully gutted Ad hold great promises as gene transfer vector for treating genetic diseases such as Duchenne muscular dystrophy. HDAd are more advantageous than E1-deleted Ad (first generation) because they encode either no or few viral genes and are consequently less immunogenic. However, in comparison to first generation Ad, the level of transgene expression was significantly weaker after gene transfer with a fully deleted HDAd in muscle even if the transgenes were regulated by the same cytomegalovirus (CMV) promoter (Gilbert et al., Hum. Gene Ther. 12:17411). In an attempt to improve the transgene expression, we have tested a partially gutted HDAd (AdRP1001) that encodes the E1B and E4 regions of Ad genome and b-galactosidase (b-gal) as transgene controlled by the CMV promoter.
DESIGN/METHODS: AdRP1001 was propagated on 293Cre cells and purified by centrifugation on CsCl gradients. The tibialis anterior (TA) muscles of neonatal, dystrophin-deficient mdx mice, Cd1 and immunodeficient scid mice were injected with 5 ml of AdRP1001 at a titer of 1.0 X 10
12 virus/ml and the number of muscle fibers expressing b-gal was analyzed at 10, 30, 90 and 180 days later by histochemistry. The TA of neonatal mdx mice was also injected with 5 ml or HDCBDys alone or in the presence of AdRP1001, both vectors being injected at the same titer (5.0 X 1011 virus/ml). The number of dystrophin positive fibers was then evaluated 10 days later by immunohistochemistry. HDCBDys is a HDAd encoding the full-length dystrophin regulated by the strong hybrid CMV enhancer/b-actin promoter (CB) promoter.
RESULTS: At 10 days post-injection, the average number of b-gal positive fibers was 792 (47% of total), 1113 (65% of total) and 820 (48% of total) in mdx, Cd1 and scid muscles respectively. Such a high transduction level has not been achieved previously by us using a fully deleted HDAd encoding dystrophin regulated by the CMV promoter (Gilbert et al., Hum. Gene Ther. 12;1741). Furthermore, no reduction in the number of transduced fibers was observed at 30, 90 and 180 days post-injection in scid mice. In mdx muscles, an average of 38 dystrophin positive fibers was observed after gene transfer with HDCBDys alone. This number increased to 378 in the presence of AdRP1001.
CONCLUSIONS: Ad gene products (E1B and/or E4) encoded by AdRP1001 can dramatically improve the expression of a transgene carried by a gutted Ad in muscle when the transgene is controlled either by the CMV or the CB promoter. More importantly, this elevated early transduction level remained unabated for at least 180 days in immunodeficient animals. The inclusion of E1B and/or E4 in a gutted Ad encoding dystrophin will most likely help in obtaining elevated and prolonged dystrophin expression in dystrophin-deficient muscle.
Supported By: Muscular dystrophy association of USA and Canada
 

[P01.064] Therapeutic Gene Replacement to Skeletal Muscle Made Efficient without Viral Vectors
George Karpati, Renald Gilbert, An-Bang Liu, Joon-Shik Moon, Basil J. Petrof, Josephine Nalbantoglu Montreal, QC, Canada; Hualien, Taiwan; Monteal, QC, Canada

OBJECTIVE: To evaluate if intramuscular injection of naked DNA (plasmid), followed by percutaneous application of electric current (electroporation, EP) is a potentially useful strategy for treating genetic diseases such as Duchenne muscular dystrophy.
BACKGROUND: Plasmid is an alternative to viral vectors for the transfer of therapeutic genes into muscle. Plasmids are easier to produce, safer and cheaper then viral vectors. However, transfection level after intramuscular injection of plasmid is very low. EP has been demonstrated to greatly improve the transfection level after plasmid injection into muscle. Pre-treatment of muscle with hyaluronidase (Hy) increases the transfection level after EP. In the present study, we have investigated the effects of age and strain of animals on EP and Hy. Two age groups and three strains of mice were studied after intramuscular injection of plasmids.
DESIGN/METHODS: Three plasmids were used: pCBLacZ and pCBMuDys encoding b-gal and murine dystrophin respectively, both transgenes being controlled by the hybrid cytomegalovirus (CMV) enchancer/b-actin promoter (CB promoter); plasmid pCMVUtrFl encoding murine utrophin regulated by the CMV promoter and tagged with a Flag epitope. The TA of 5-week-old Cd1, C57BL/6 and dystrophin-deficient mdx mice, as well as 15-day-old mdx mice were injected with 1 mg/ml of plasmid followed by 8 pulses (duration: 20 msec; interval: 1 sec) of 175-200 V/cm. Some TA were also injected with Hy (0.4m/ml) two hours before EP. At 6 to 8 days post-injection, the transfection level was determined by processing the muscles for histochemistry and immunohistochemistry.
RESULTS: In adult mdx, C57Bl/6 and Cd1 animals, plasmid alone produced 3-10 transfected fibers. In the same model, EP markedly increased the transfection level of b-gal expressing plasmid in TA muscle to 169 (10%), 279 (16%) and 326 (19%) fibers respectively. EP+Hy further increased the transfection level by 2-3 folds. Although a trend was observed toward lower transduction level in mdx mice, the difference was not significant. In very young mdx animals, b-gal expressing plasmid plus EP produced very low transfection (5 fibers/muscle). EP+Hy produced 120 fibers/muscle. Adult mdx muscles injected with plasmid expressing either murine dystrophin or utrophin plus EP+Hy produced up to 144 and 276 transfected fibers/muscle respectively.
CONCLUSIONS: Pre-treatment with Hy followed by EP markedly increase the transduction level after intramuscular injection of plasmid DNA in adult normal (Cd1 and C57Bl/6) and dystrophic muscles (mdx). The relatively high level of dystrophin and utrophin gene transfer following Hy and EP treatments will permit to study in dystrophic muscles many aspects of dystrophin vs utrophin gene transfer without interference by viral vectors.
Supported By: Muscular dystrophy association of USA and Canada.
 

P01.067] Novel Dystrophin Gene Mutations and Unusual DMD/BMD Phenotypes
Elicia Estrella, Benjamin Roa, Thomas Prior, Jerry Mendell, Lisa Baumbach Miami, FL; Houston, TX; Columbus , OH

OBJECTIVE: This study illustrates several unusual clinical presentations of DMD/BMD.
BACKGROUND: Since the discovery of the Duchenne Muscular Dystrophy (DMD) gene approximately 15 years ago, hundreds of Duchenne, Becker, and other patients with a suspected primary dystrophinopathy have been evaluated for mutations in the DMD/BMD locus. Knowledge gained from these studies has led to general genotype: phenotype predictions, although these predictions do not nearly account for the spectrum of clinical variability noted. The detection of patients with novel mutations may provide new insights into functional aspects of the dystrophin gene and/or associations with unusual clinical features.
DESIGN/METHODS: Patients were evaluated for deletions/ duplications in the dystrophin gene. Molecular analyses were completed via standard PCR and southern blotting techniques. In several patients, Western blot analysis was performed for quantitative and qualitative studies.
RESULTS: We report our investigations of several unusual DMD/BMD patients. The first two patients displayed no previous family history of muscle disease and/or similar symptoms. Patient 1, an 11 1/2 year old Caucasian male, was referred for a long undefined history of weakness and unsteadiness. He had moderate serum CK elevations. At time of examination, he had no pseudohypertrophy, no significant weakness, a modified Gowers maneuver, and normal reflexes in all extremities. His only clinical complaint was of pain (cramps) after strenuous exercise. Mutation studies of the DMD gene revealed a deletion spanning exons 3-4. This deletion has been reported in three other Beckers patients (Leiden Muscular Dystrophy mutation database). Although the deletion is found in the beginning of the gene, and presumably affects mRNA splicing, it is very interesting that all patients exhibit an extremely mild Beckers phenotype. Patient 2, a 6 1/2 year old African-American male, displays marked developmental delay for all milestones (motor and cognitive) since infancy. He also has speech delay, and ADHD. These features are not present in any other family members. Clinical exam noted minimal muscle involvement including upper extremity weakness (since age 3), mild calf pseudohypertrophy, no ambulatory problems, but markedly elevated serum CK levels. DNA studies revealed a possible duplication of exons 10-12 and the African-American exon 8/9 RFLP. Review of the literature indicates that this is a novel mutation, associated with a gross gene rearrangement. His phenotype is unusual for the marked level of mental impairment. Patients three-five are all classical BMD patients who share a common exon 45-53 in-frame deletion. Western blot analysis of muscle dystrophin revealed a normal level (100%) of the truncated, mutant form. DMD/BMD deletions which produce quantitatively normal dystrophin levels should be further investigated.
CONCLUSIONS: It is our hope that similar patient descriptions will lead to further understanding of the dystrophin-associated muscular dystrophies. Insight gained from these genotype:phenotype correlations may lead to better progostic and treatment capabilities.
 

[P04.145] Relatively Prevalent Mutations in the Dysferlin Gene and Genotype-Phenotype Correlation of Miyoshi Myopathy in Japanese Population
Toshiaki Takahashi, Masashi Aoki, Maki Tatayama, Yoshiaki Onodera, Yuji Hinuma, Emi Kondo, Ikuya Nonaka, Kiichi Arahata, Robert H. Brown, Jr., Hiroshi Saito, Yasuto Itoyama Sendai, Miyagi, Japan; Kodaira, Tokyo, Japan; Charlestown, MA

OBJECTIVE: To reveal the features of dysferlin gene mutations in Japanese patients with Miyoshi myopathy (MM) and a genotype-phenotype correlation in the disease.
BACKGROUND: MM is an autosomal recessive distal muscular dystrophy. The gene dysferlin was identified recently and found to be mutated both in families with MM and those with limb girdle muscular dystrophy type 2B.
DESIGN/METHODS: We examined 26 Japanese patients with MM. Genomic DNA was extracted from the peripheral lymphocytes of the patients with informed consent. The PCR products of each 55 exon were screened by single strand conformation polymorphism (SSCP) or direct sequencing from the PCR fragments.
RESULTS: We identified 16 different mutations in 20 MM patients and 10 of them were novel. Mutations in Japanese patients are distributed along the entire length of the gene. The mean age at onset of the patients with the dysferlin mutations was 23.7 ± 7.3 years (from 14 years to 37 years). The mean serum CK level of the patients was 5,326 ± 3,152 IU/l (min 1,289 IU/l, max 12,556 IU/l). Residence of the patients with the dysferlin mutations had a wide distribution in Japan.
CONCLUSIONS: We found relatively prevalent four mutations (C1939G, G3370T, 3746delG, and 4870delT) in these populations, and they accounted for 60 percent of the mutations in this study. We speculated that most patients with MM were selectively affected in the paravertebral muscles even in the very early stage. This study revealed that the G3370T mutation may be associated with the mild form of MM whereas G3510A mutation may be associated with the severe form of MM.
Supported By: Grant-in-Aid from the Ministry of Health, Labor and Welfare in Japan and Grant-in-Aid for Scientific Research from Japan Society for the Promotion of Science.
 

P01.063] Myogenesis of Human Adult Dermal Fibroblasts (HADFs) In Vitro:: Potential Source for Gene Therapy of Muscular Dystrophies
Tokuro Furuya, Hiroaki Tanaka, Noriyoshi Kameda, Fumiko Saito, Takayoshi Kobayashi, Hidehiro Mizusawa Bunkyo-ku, Tokyo, Japan; Nakano-ku, Tokyo, Japan

OBJECTIVE: To clarify whether HADFs particitate in the myotube formation.
BACKGROUND: Skeletal muscles are generated mainly from myogenic precursor cells. However, recently some reports have indicated that other type of cells could become the source of myogenesis such as bone marrow.
DESIGN/METHODS: HADFs from human skin biopsied with informed consents were cultured in several passages and labeled with marker gene LacZ encoding bacterial beta-galactosidase. Same numbers of labelled HADFs and mouse myoblasts, C2C12 cells, were co-cultured. To identify the origin of the nuclei of myotubes in co-culture, we performed Fluorescence in situ hybridization (FISH) using two probe which contain human or mouse specific chromosome repetitive sequence. To investigate whether HADFs-nuclei in myotubes expressed muscle specific transcripts, RT-PCR primers were designed in the 3' UTR region of human myogenin gene. HADFS with marker gene were co-cultured with human muscle cells innervated with fetal rat spinal cords whether they particitate mature innervated muscle fibers.
RESULTS: When HADFs were co-cultured with mouse myoblasts, beta-galactosidase positive myotubes appeared, which were supposed to have HADFs-derived nuclei. FISH revealed that these myotubes have at least one human nuclei among their multiple nuclei. In co-culture of HADFs and mouse myoblasts, human specific myogenin transcripts were detected by RT-PCR. Furthermore when LacZ introduced HADFs were co-cultured with human muscle cells innervated with fetal rat spinal cord, beta-galactosidase positive striated contracting mature muscle fibers were formed.
CONCLUSIONS: This is the first report to clarify that HADFs have the potential capacity of participating myogenesis in vitro. HADFs may become the alternative source of cell transplantation therapy for muscular dystrophies

[P01.066] Effect of Daily Prednisone on Independent Ambulation in Patients with Duchenne Dystrophy Treated for up to 15 Years
Shree Pandya, Gary J. Myers, Richard T. Moxley Rocheste, NY; Rochester, NY

OBJECTIVE: To document the effect of long term prednisone treatment on ambulation in patients with Duchenne dystrophy.
BACKGROUND: Three double blind, randomized, controlled trials and a three year follow up open trial have clearly documented that prednisone increases muscle strength and muscle mass, and slows progression of disease. Prednisone administered daily at a dose of 0.75mg/kg/d increases strength over a three month period, stabilizes it for eighteen months or more, and slows progression of the disease over at least a three year period. DeSilva et al in a retrospective study of 16 patients found that a subset of patients who received prednisone for 2 years or more, lost independent ambulation at an average age of 13.1, compared to the control group who lost independent ambulation at 10.3, a gain of almost 3 years.
DESIGN/METHODS: We have prospectively followed a group of patients with Duchenne dystrophy who began prednisone treatment between 1986 and 1989 as part of the Clinical Investigation of Duchenne Dystrophy (CIDD) therapeutic trials at our center. After completion of the trials, these patients have received 0.75mg/kg/d of prednisone unless side effects required a reduction in dosage. Patients have returned twice yearly for evaluations which included measurements of vital signs, height, weight, timed function tests and side effects.
RESULTS: We have followed 30 patients for an average of 11.2 years (range 3-15 yrs). At the initiation of prednisone, 18 of the 30 were ambulatory. 13 walked independently (avg age 8.3 yrs) and 5 walked with long leg braces (avg age 10 yrs). At last contact all patients had become wheelchair dependent. One patient has been lost to follow up. Three patients discontinued Prednisone due to weight gain. The average age at loss of independent ambulation is 14.6 years (range 12 to 19 yrs).
CONCLUSIONS: Loss of independent ambulation has been reported to occur between the ages of 8.5 and 10.8 years in untreated patients. In our series, patients treated with prednisone have ambulated independently until age 14.6, a gain of 4 to 6 years compared to untreated controls reported in the literature. Long term treatment with prednisone substantially lengthens independent ambulation in patients with Duchenne dystrophy.

S24.005] Dystrophin Gene Repair in mdx Muscle Precursor Cells Mediated by RNA/DNA Chimeric Oligonucleotides
Thomas A. Rando, Carmen Bertoni Stanford, CA

OBJECTIVE: To test for chimeraplast-mediated repair of the dystrophin gene point mutation in mdx muscle precursor cells.
BACKGROUND: Chimeraplasts are chimeric oligonucleotides made up of both DNA and RNA bases that have been found to be effective as gene therapy agents to correct point mutations in genomic DNA. They are designed to be homologous to a region of DNA with a single mismatch at the base targeted for correction. The mechanism of repair involves first homologous pairing followed by the induction of cellular mismatch repair activities. Chimeraplasts have been shown to induce repair of point mutations in different cell types, both in vivo and in vitro. We have previously demonstrated that injection of MDX1, a chimeraplast targeted to correct the point mutation in the mdx dystrophin gene, into muscles of mdx mice leads to repair of that mutation and to dystrophin protein expression in differentiated myofibers in vivo. We wanted to determine whether MDX1 would also correct the point mutation in mdx muscle precursor cells (satellite cells or myoblasts) in order to further assess the potential for chimeraplasts to be developed for in vivo or ex vivo gene therapy.
DESIGN/METHODS: Myoblasts were derived from mdx muscle and propagated in vitro. MDX1 or control (MDX2) chimeraplasts were transfected into those cells. After growth in culture, repair of the mdx point mutation was assessed at the genomic level using the mdx-ARMS assay. Alternatively, cells were induced to differentiate into myotubes, and correction of the mutation was assessed at the mRNA level by RT-PCR-ARMS assay. Differentiated cultures were also assessed for dystrophin protein expression using both Western analysis and immunohistochemical staining. For in vivo experiments, chimeraplasts were directly injected into muscles of mdx mice. Two days later, primary cultures were established from those injected muscles and the myoblasts in the cultures were assessed for dystrophin gene correction by the same methodologies. Wild-type (C57) myoblasts and mice were used as controls.
RESULTS: Correction of the mdx point mutation by MDX1, but not MDX2, was demonstrated at the genomic level in muscle precursor cells after in vitro transfection or in vivo injection. Similarly, transcripts containing the wild-type dystrophin sequence were demonstrated in mdx myotube cultures in both experimental paradigms. Immunohistochemical analysis of differentiated cultures demonstrated dystrophin protein expression in multinucleated myotubes. Western analysis demonstrated that the level of dystrophin expression in MDX1-transfected cells, 4 weeks after transfection and 3 days after differentiation, ranged from 2% to 15% (n=6) of that seen in C57 myotube cultures.
CONCLUSIONS: Chimeraplasts can effect gene repair in muscle precursor cells, both in vitro and in vivo. This is a promising finding for using chimeraplasts for either in vivo or ex vivo gene therapy for dystrophies due to point mutations of the dystrophin gene. Further studies are focusing on methods to improve the efficiency of gene repair still further and establishing methods for in vivo delivery of chimeraplasts.
Supported By: Study supported by a grant from the Muscular Dystrophy Association to TAR.

P01.068] Human Myogenic Precursor Cells (mpc) Attract Monocytes through CC and CX3C Chemokines and Interplay with Macrophages To Improve Muscle Regeneration: A Step toward Characterization of the Mpc Niche
Romain K. Gherardi, Corinne Sonnet, Peggy Lafuste, Benedicte Chazaud Creteil, France, Metropolitan

OBJECTIVE: To characterize the interplay between myogenic precursor cells (mpc) and monocyte/macrophages (m/M).
BACKGROUND: Cell therapy using myoblast transfer is based on capacity of mpc to be activated after muscle injury, to proliferate and to fuse into new muscle fibers. To develop their myogenic program, mpc depend from environmental cues, the so-called biologic niche. Macrophages are crucially involved in skeletal muscle regeneration: after injury, the muscle fiber is invaded by macrophages and regeneration is markedly impaired in the absence of macrophage infiltration. In addition to muscle cell debris removal by phagocytosis, macrophages likely participate to the subsequent phases of muscle reconstruction. Investigating this topic should allow optimisation of future cell therapy programs.
DESIGN/METHODS: Mpc:m/M interplay was assessed using human cells according to a 3-step methodology: 1) assessment of elementary biologic effects on cell density, phagocytosis, and chemotaxis through porous filters or endothelial cells; 2) selection of candidate effector molecules for the observed effects using a DNA array procedure; 3) validation of the role of selected candidate molecules by specific inhibition tests.
RESULTS: Mpc exerted a chemotactic effect toward monocytes, which effect was potentiated by 35% when mpc were previously stimulated by macrophages. Macrophages alone exerted a weaker chemotactic effect toward monocytes than mpc alone, but stimulation of macrophages by mpc increased the chemotactic activity of macrophages by 80%.
DNA array displayed expression of three chemokines by mpc that could be responsible for chemotaxis: MCP-1 (CCL2), MDC (CCL22) and fractalkine (CX3CL1). Monocyte chemotaxis toward mpc was inhibited from 40 to 60% in the presence of blocking antibodies against these chemokines. The 3 antibodies mixed together reduced chemotaxis by 90%.
Mpc:M cocultures showed no phagocytosis of living mpc. Moreover, macrophages increased mpc density in a dose-dependent manner, a 1:2 (mpc:M) ratio inducing a 85% increase of mpc density.
CONCLUSIONS: Activated human mpc recruit circulating monocytes and stimulate the recruited macrophage chemotactic activity. In turn, macrophages increase mpc chemotactic activity and boost mpc growth.
Supported By: Association Francaise contre les Myopathies (AFM)

[S24.002] RNA Splicing Defect Leads to Chloride Channelopathy and Myotonia in Myotonic Dystrophy (DM)
Ami Mankodi, Christina Jiang, Masanori P. Takahashi, Carol Beck, Richard T. Moxley, Stephen C. Cannon, Charles A. Thornton Rochester, NY; Boston, MA; Philadelphia, PA; Boston, MA

OBJECTIVE: Determine the cause of myotonia in DM type 1 (DM1) and DM type 2 (DM2).
BACKGROUND: DM is caused by expansion of a CTG repeat on chromosome 19 (DM1) or a CCTG repeat on chromosome 3 (DM2). Manifestations of DM result from accumulation in the nucleus of RNA with expanded CUG repeats (DM1) or CCUG repeats (DM2). Transgenic mice that express expanded CUG repeats (HSA-LR mice) also develop myotonia and a DM-like myopathy. While myotonia implies dysfunction of ion channels, the nature of the channelopathy in DM is unknown.
DESIGN/METHODS: Ion conductance in the transgenic model was assessed by making intracellular recordings from muscle fibers of HSA-LR or wild-type (WT) mice. Expression of the chloride channel 1 (ClC-1) mRNA in murine and human DM was assessed by Northern blot, real-time RT-PCR, and sequence analysis of cDNA clones to evaluate RNA splicing. Distribution of Cl channel was assessed by immunofluorescence (IF) using polyclonal antibody to the C-terminus of ClC-1.
RESULTS: Compared to WT mice, Cl conductance in HSA-LR mice was reduced 10-fold, an alteration sufficient to cause myotonia. To investigate the mechanism for reduced chloride conductance, we examined the expression of ClC-1, the predominant Cl channel in muscle. HSA-LR mice showed a major defect in splicing of the ClC-1 pre-mRNA. Among 29 ClC-1 cDNA clones, only 24% encoded full-length protein. 76% of clones were incorrectly spliced; nearly all had premature termination codons. A wide range of splicing errors was observed (11 different anomalous splice sites). The total level of ClC-1 mRNA in HSA-LR muscle was slightly reduced (~40% of WT), presumably due to accelerated degradation, via the nonsense-mediated decay pathway, of mis-spliced mRNA. Results in human DM were similar: the fraction of aberrantly-spliced ClC-1 clones was >95% in severely-affected patients with DM1 (n=2), and > 70% in patients with DM2 (n=2). IF on muscle sections from HSA-LR mice and from patients with DM1 or DM2 revealed a mosaic pattern in which some muscle fibers had a normal circumferential rim of ClC-1 protein, but most were devoid of ClC-1 or had interruptions in the sarcolemmal rim of staining.
CONCLUSIONS: Our results support a model in which nuclear accumulation of RNA with expanded CUG or CCUG repeats causes aberrant splicing of ClC-1 mRNA, loss of ClC-1 protein, reduced chloride conductance, and consequently hyperexcitability of the muscle membrane in both DM1 and DM2.
Supported By: Supported by NIH and the Muscular Dystrophy Association.

 P04.144] A Novel, Blood-Based Diagnostic Assay for Dysferlin Myopathies(Limb Girdle Muscular Dystrophy 2B, Miyoshi Myopathy and Distal Anterior Compartment Myopathy)
Eduard Gallardo, Mengfatt Ho, De Luna Noemi, Diane Mckenna-Yasek, Robert H. Brown, Isabel Illa Barcelona, Spain; Boston, MA

OBJECTIVE: To develop a new dagnostic assay for limb girdle muscle dystrophy 2B (LGMD2B), Miyoshi myopathy (MM)and distal anterior compartment myopathy (DAT), which screens for dysferlin expression in blood using a commercially available monoclonal antibody.
BACKGROUND: Recently, LGMD2B,MM and DAT were found to be allelic disorders arising from defects in a novel gene that encodes dysferlin. The dysferlin gene is large comprising 55 exons and encodes a protein of 237 kDa. Dysferlin is localized to the plasma membrane of muscle fibers. Its function is unknown, however its homology with a gene of the nematode Caenorhabditis elegans,fer-1, suggests that it may function in membrane fusion or trafficking. The size of the gene and the fact that most mutations are single nucleotide changes make molecular diagnosis difficult. Western Blot(W-B) analysis of muscle biopsies has been shown a reliable and rapid method for the diagnosis of muscle dystrophies. However, muscle biopsy is an invasive procedure.
DESIGN/METHODS: We isolated peripheral blood mononuclear cells (PBMC) from whole blood of patients with disferlinopathy (n=12) and controls using a Ficoll-Plaque gradient centrifugation. Total RNA was isolated and we performed a Northern-Blot (N-B)with specific probes. W-B and immunocytochemistry using a monoclonal antibody anti-dysferlin (NCL-Hamlet)were also performed. In addition, we isolated CD14+ (a monocyte marker) cells by means of magnetic beads and column separation, and performed W-B and immunocytochemistry using the same antibody.
RESULTS: N-B analysis of total RNA from blood showed a weak but distinct band of about 7.5 kb, corresponding to the dysferlin transcript. A prominent band of 230kDa was detected by W-B using PBMC extracts from controls but not in patients. Immunocytochemistry confirmed the results. Immunocytochemistry and W-B of purified CD14+ cells verified that expression of dysferlin was restricted to monocytes. Skeletal muscle from all patients was also tested for dysferlin expression and it was absent in all of them.
CONCLUSIONS: Dysferlin is expressed in monocytes CD14+ and correlates with its expression in skeletal muscle. This new blood-based diagnostic assay is a simple and rapid technique and offers a less invasive tissue sampling compared to muscle biopsy. The discovery of expression in monocytes provides a new paradigm to study the biological function of this protein in a non-muscle cell type that is more accesible. Also, for experimental therapies intended to increase the levels of dysferlin, we could now track efficacy by monitoring dysferlin levels in blood rather than skeletal muscle.
Supported By: FIS 99/019-01; FIS 01/0979; MDA (USA); C.B.Day Investment Company; NIH

[
P03.126] Creatine Monohydrate in Proximal Myotonic Myopathy (PROMM): A Double Blind Placebo Controlled Clinical Study
Christiane Schneider, Marcus Beck, Annette George, Carsten Wessig, Henrik Kele, Wolfram Kress, Karlheinz Reiners, Klaus V. Toyka Würzburg, Germany; Göttingen, Germany

OBJECTIVE: To investigate the efficacy and safety of creatine monohydrate (Cr) in patients with the proximal myotonic myopathy (PROMM) phenotype.
BACKGROUND: PROMM is characterised by proximal weakness, myotonia, myalgia and cataracts. Fluctuations of weakness, myotonia and muscle pain occur frequently in PROMM suggesting that disturbances of muscle energy metabolism might be involved. Cr is known to have various positive effects on muscle cell metabolism and was shown to be efficient in muscular dystrophies and mitochondrial myopathies.
DESIGN/METHODS: Twenty patients (10 men, 10 women) with PROMM with overt neuromuscular weakness and myotonia were randomised to either the Cr or the placebo group. Cr was given for 3 months in a daily dose of 10 g, i.e. 18 ml of pure Cr powder. Controls received the same volume of microcrystalline cellulose. muscle strength sum scores using the Medical Research Council (MRC) scale, the Neuromuscular Symptom Score (NSS) and hand-held dynamometry, and a self assessment analogue scale were evaluated seperately for each treatment group before and after the treatment period by a blinded examiner. We defined changes in average MRC and hand-held dynamometry sum scores and NSS as primary endpoints.
RESULTS: Using the strength measurement methods described above no significant improvement of muscle strength was found in the total Cr group as compared to the placebo group and in the male patients of both groups (Mann-Whitney U-test, p > 0.05). In the gender subgroup analysis muscle strength improved significantly in women who had received Cr as compared to women who had received placebo (p = 0.005 (hand-held dynamometry) and p = 0.03 (MRC scale), Mann-Whitney U-test). Seven patients of the Cr group and one patient of the placebo group reported subjective improvement with increased endurance in daily activities and sports. During the administration of Cr leg muscle pain resolved only in one male and one female patient. Chest pain resolved in another male patient.
CONCLUSIONS: Cr was well tolerated during the treatment period. In this pilot study, Cr had a beneficial effect on muscle strength only in women. In addition, Cr occasionally had a positive effect on muscle and chest pain. Larger studies are necessary to confirm the results of this study.
Supported By: Verein der Freunde und Förderer der Neurologischen Klinik der Universität Würzburg. Cr and placebo medication were kindly provided by SKW Trostberg, Trostberg, Germany.

[P03.132] Cardiac Involvement in Young Myotonic Dystrophy (DM1) Patients
Guillaume Bassez, Isabelle Desguerre, Arnaud Lazarus, Pascal Laforêt, Henri Marc Bécane, Jean Varin, Hélène Radvanyi, Bruno Eymard, Denis Duboc Créteil, France; Paris, France

OBJECTIVE: To determine the frequency of severe cardiac symptoms in young patients with myotonic dystrophy (DM1).
BACKGROUND: Cardiac symptoms, one component of DM1 systemic features, have been identified as a crucial prognostic factor in adults. Significant early cardiac involvement in young affected individuals is usually considered as uncommon. However, there is little available data concerning cardiac symptoms for children and young adults. It is also unclear whether severe cardiac signs may be encountered as an isolated inaugural symptom in DM1 adolescents, or constantly combined with other significant clinical DM1 signs. Cardiac evaluation in young DM1 patients (<18 year old) may be of great interest for both follow-up and management of cardiac disorders in childhood type of DM1, and may add useful information for genetic counselling for asymptomatic individuals at risk under the age of 18.
DESIGN/METHODS: In a retrospective study of DM1 patients, we analysed cases with severe cardiac involvement identified before the age of 18. Age at onset, detailed cardiac, neuromuscular, and other systemic clinical features, results of cardiac investigations (conduction defects, arrhythmia, ventricular dysfunction), severity, follow-up, and number of (CTG)n triplet repeats were investigated.
RESULTS: Six young individuals were identified with severe cardiac involvement before the age of 18. Two subgroups can be differentiated : cardiac signs during previously diagnosed childhood or congenital type of DM1, and severe cardiac events inaugurating DM1 in asymptomatic teenagers. In the first group (n=3), cardiac abnormalities were sudden death in a 17 year old boy with pace-maker, due to ventricular tachycardia after exercise; recurrent ventricular tachycardia episodes during dancing in a 12 year old girl; and a worsening intraventricular conduction defect in a 15 year old girl, requiring pace-maker at the age of 22. In the second group (n=3), a 16 year old asymptomatic boy presented with cardiac arrest after running with no recurrence after implantable cardiac defibrillator; a 14 year old boy with mild distal muscle weakness had atrial fibrillation; and recurrent pre-syncope events due to paroxysmal atrial flutter were diagnosed in a clinically unaffected 13 year old girl. A wide variation of CTG expansion was observed (from 250 to 1200 repeats).
CONCLUSIONS: Severe cardiac symptoms may occur early in the course of DM1 in two different manners. At various ages in congenital/childhood type affected patients, as well as in clinically asymptomatic adolescents with minor muscle signs. Moreover, cardiac manifestation may be the first isolated clinical sign of DM1. No particular range of CTG expansion appear predictive. In addition to our data, further prospective studies may be useful for accurate cardiac management during infancy/adolescence as well as for genetic counselling in young DM1 at risk individuals.
Supported By: Association Francaise contre les Myopathies (AFM)

[P03.130] Clinical and Molecular Correlations in Myotonic Dystrophy Type 2
John W. Day, Kenneth Ricker, Katherine A. Dick, Laura Rasmussen, Joline C. Dalton, Jennifer F. Jacobsen, Christina L. Liquori, Wolfram Kress, Christiane Schneider, Melinda L. Moseley, Laura P. W. Ranum Minneapolis, MN; Wurzburg, Germany

OBJECTIVE: To determine the breadth of clinical features in myotonic dystrophy type 2, and to determine the correlation of those features with the size of the tetranucleotide CCTG repeat expansion responsible for the disease.
BACKGROUND: Myotonic dystrophy (DM) causes a distinctive constellation of seemingly unrelated clinical features that can be caused by mutations on either chromosome 19 (DM1) or 3 (DM2). We recently demonstrated that a CCTG expansion in intron 1 of ZNF9 causes DM2. We now report the clinical and molecular correlations of this mutation in approximately 250 affected individuals from 82 families.
DESIGN/METHODS: Neurological and electrophysiological examinations of many patients were carried out in the field, while others were studied in more detail in clinic.
RESULTS: Individuals affected by DM2 have a complex clinical presentation that is strikingly similar to DM1: a characteristic pattern of weakness involving facial, neck flexor, finger flexor and hip girdle musculature; iridescent cataracts that can be symptomatic in the 3rd decade of life; hypotestosteronism and oligospermia in males; hypogammaglobulinemia, insulin insensitivity, and elevated creatine kinase; potentially fatal cardiac involvement involving severe arrhythmias or progressive cardiomyopathy. As in DM1, skeletal muscle biopsies show the characteristic features of severe fiber atrophy and a profusion of central nuclei, with intranuclear RNA inclusions in essentially all nuclei. We investigated various aspects of disease severity and found no correlation with CCTG repeat length. To date there are no known cases of severe congenital onset of DM2, and the cognitive involvement appears to be much less severe than can occur in DM1.
CONCLUSIONS: The DM1 CTG and DM2 CCTG expansions cause a strikingly similar constellation of clinical features, electrophysiological abnormalities and muscle histopathology including the presence of RNA foci. The clinical similarities of DM1 and DM2 indicate that the downstream effects of microsattelite expansions expressed in RNA cause the features common to both disesases.
Supported By: NIH/NINDS NS35870-05
NIH/Univ MN GCRC
MDA

[P03.128] The Prevalence of Myotonic Dystrophy in Israeli Jewish Communities: Inter-Community Variation and Founder Premutations
Rival Segel, S. Silverstein, I. Lerer, E. Kahana, R. Meir, M. Sagi, N. Zilber, Amos D. Korczyn, Y. Shapira, Z. Argov, R. Carmi, Zipora Falik, C. Ligum, Moshe Frydman, Z. Gelman-Kahan, S. Shalev, Y. Finkelstein, Devora Abeliovich Jerusalem, Israel; Israel; Tel-Aviv, Israel

OBJECTIVE: We attempted to perform a comprehensive epidemiological and genetic survey of MyD among Jews living in Israel.
BACKGROUND: Myotonc dystrophy (MyD) occurs all over the world but exact data on prevalence and frequency of premutations of the unstable DMPK-(CTG)n allele on chromosone 19 in various populations are lacking.
DESIGN/METHODS: The frequency of unrelated sibships in each community was used as an estimate of the transition rate from stable to unstable DMPK-(CTG)n alleles assuming that each transition founds a new MyD sibship.
RESULTS: The average prevalence of myotonic dystrophy (MyD) was 15/105 with inter-community variations; the Ashkenazi Jews had the lowest rate (5.7/105) which was much higher among Yemenite Jews (47.3/105; p<0.05). This study indicated that the differences in the prevalence of MyD are a result of higher mutation rate in the non-Ashkenazi Jews (>50/108) as compared to the rate in the Ahskenazi Jews (16.3/108). The intragenic haplotypes of the MyD alleles were the same as in many populations over the world. However, two MyD closely linked markers D19S207 and D19S112 were in linkage dysequilibrium with the MyD mutation in patients of Yemenite and Moroccan extractions but not in Ashkenazi patients. This observation indicated a common ancestral origin for the MyD premutation in patients of a given ethnic origin.
CONCLUSIONS: The difference in the prevalence of MyD among Jewish communities is a consequence of high frequency of founder premutations among non-Ashkenazi Jews, and particularly Yemenite Jews.