RESUMOS PRINCIPAIS DO CONGRESSO ANUAL DA SOCIEDADE AMERICANA DE GENÉTICA HUMANA

ABSTRACTS - THE ANNUAL METTING OF AMERICAN SOCIETY OF HUMAN GENETICS

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Bisphosphonate treatment for osteopenia associated with genetic disease in children. S. Carter1, R. Mendoza-Londono1, V.R. Sutton1, C.A. Bacino1, K. Ellis2, R. Shypailo2, B. Lee1. 1) Molecular and Human Genetics, Baylor College of Medicine, Houston, TX; 2) Dept. Pediatrics, CNRC, Baylor College of Medicine.

   Osteoporosis is defined as low bone mass and bone fragility with increased susceptibility to fractures. It is also defined as bone mineral density (BMD) that is 2.5 SD below the mean peak value for age-matched controls. Until recently, treatment options for pediatric osteoporosis have been limited. Bisphosphonates inhibit osteoclast activity and have been shown to be safe in pediatric applications. Common genetic conditions associated with low BMD and susceptibility to fractures include osteogenesis imperfecta (OI), and secondary osteopenia due to to lack of physical activity, as is seen in patients with neuromuscular disease. We present our experience treating 14 pediatric patients followed at the Texas Childrens Hospital Skeletal Dysplasia Clinic between 1999 and 2002. The diagnosis was OI type 3 or 4 in 11 patients and other neuromuscular disease in 3 patients. Ages ranged between newborn and 15 yrs. (Average 4.5 yrs). The diagnosis of OI was made in the first two months of life in 9 of the 11 patients. Bone mineral density of the lumbar spine, femoral head and total body was measured using dual-energy x-ray absorptiometry (DXA) before treatment and at regular intervals during treatment. The patients received Pamidronate at a dose of 13.5 mg/kg/year for children <2-3 years and 9 mg/kg/year for children >3 years. Bone metabolic parameters (alkaline phosphatase, urine collagen N-telopeptides) showed decrease in bone turnover. Average L-spine BMD values were Z= -3.86 (Range 6.78 to 2.92) before treatment and Z= -2.56 (range 3.72 to 1.4) after treatment. Physiologic response appeared more significant in younger children. Overall there was a subjective clinical improvement, with decrease in bone pain and frequency of fractures, resulting in better tolerance of daily activities and well being. Side effects were minor including transient febrile episodes and mild hypocalcemia which corrected with supplementation. This experience has demonstrated that bisphosphonates constitute an important adjuvant treatment for osteoporosis associated with genetic disease.

Targeted exon skipping as a promising therapeutic tool for Duchenne muscular dystrophy. J.C.T. Van Deutekom1, A. Aartsma-Rus1, M. Bremmer-Bout1, J.A.M. Janson1, W.E. Kaman1, E.J De Meijer1, F. Baas2, J.T. Den Dunnen1, G.J.B. Van Ommen1. 1) Dept Human Genetics, Leiden Univ Medical Ctr, Leiden, Netherlands; 2) Dept Neurology, Academic Medical Ctr, Amsterdam, Netherlands.

   Duchenne muscular dystrophy (DMD) is a lethal muscle disease typically caused by frame-shifting mutations in the DMD gene that abort the synthesis of the dystrophin protein. In contrast to the gene therapy studies that are based on gene replacement, we are focusing on gene correction through the targeted modulation of the splicing of the patient-specific gene. Using antisense oligoribonucleotides (AONs) directed to exon-internal sequences, we aim to induce the specific skipping of a target exon in order to generate a shorter, but in-frame transcript similar to those found in the corresponding Becker muscular dystrophy (BMD) patients, having milder phenotypes and longer life expectancies. We have recently demonstrated the feasibility of AON-induced skipping of 15 different exons within the deletion hot spot regions. In this study we show the broad therapeutic applicability of skipping these exons in cultured muscle cells from DMD-patients affected by different mutations. In fact, through the AON-mediated production of novel, in-frame transcripts, we detected dystrophin synthesis in at least 70% of treated muscle cells. We here also show the in vivo feasibility of targeted exon skipping in mouse muscle tissue. Following intramuscular injections of mouse-specific exon 46 AONs, we were able to specifically induce exon 46 skipping in a dose-dependent manner. In a subsequent time-series experiment, the skipping effect was highest at 12 days post-injection with estimated efficiencies of 20% of total RT-PCR products, and, although gradually diminishing, continued for at least 29 days. Moreover, using DMD humanized transgenic mice carrying an integrated 2.7 Mb YAC-derived copy of the human DMD gene, we were able to test human sequence-specific AONs, and induce the skipping of human DMD exons, in an experimental mouse background. Our results confirm the therapeutic potential of AON-induced exon skipping, and contribute to its further development for DMD gene therapy.

a- and b-sarcoglycan delivery by recombinant adeno-associated virus: efficient rescue of muscle, but differential toxicity. D. Dressman1,2, K. Araishi3, M. Imamura3, T. Sasaoka3, L.A. Liu4, E. Engvall4, E.P. Hoffman1,2. 1) Center for Genetic Medicine, CNMC, Washington, DC; 2) University of Pittsburgh, Department of Molecular Genetics & Biochemistry, Pittsburgh, PA; 3) National Institute of Neuroscience, NCNP, Tokyo, Japan; 4) Burnham Institute, La Jolla, CA.

   The sarcoglycanopathies are a group of four autosomal recessive muscular dystrophies (LGMD 2D, 2E, 2C, and 2F), caused by mutations of the a, b, g, or d sarcoglycan genes. The d-sarcoglycan deficient hamster has been the most utilized model for gene delivery to muscle by recombinant adeno-associated virus (AAV) vectors, however human patients with d-sarcoglycan deficiency are exceedingly rare with only two patients described in the United States. Here, we report construction and use of AAV vectors expressing either a- or b- sarcoglycan, the genes responsible for the most common forms of the human sarcoglycanopathies. Both vectors showed successful short-term genetic, biochemical and histological rescue of both a- and b-sarcoglycan deficient mouse muscle. However, comparison of persistence of expression in 51 injected mice showed substantial differences between AAV a-sarcoglycan (a-SG) and b-sarcoglycan (b-SG) vectors. AAV b-SG showed long-term expression with no decrease in expression for over 21 months after injection, while AAV a-SG showed a dramatic loss of positive fibers between 28 days and 41 days post-injection (p=0.006). Loss of immuno-positive myofibers was correlated with significant inflammatory cell infiltrate, primarily macrophages. To determine if the loss of a-sarcoglycan positive fibers was due to an immune response or cytotoxic effect of a-sarcoglycan over-expression, SCID mouse muscle was assayed for cytotoxicity after injection with AAV a- SG, AAV b- SG, or PBS. The results were consistent with over-expression of a-sarcoglycan causing significant cytotoxicity. The cytotoxicity of a-sarcoglycan, and not b- or d-sarcoglycan over-expression, was consistent with biochemical studies of the hierarchical order of assembly of the sarcoglycan complex. Our data suggests that even closely related proteins might require different levels of expression to avoid toxicity and achieve long-term tissue rescue.

Immunostimulatory Properties of Dystrophic Muscle Alter Persistence of Transgenes. E.P. Hoffman1,2, H. Gordish3, K. Araishi4, M. Imamura4, T. Sasaoka4, D. Dressman1,2. 1) Research Center for Genetic Medicine, Children's National Medical Center, Washington, DC; 2) University of Pittsburgh, School of Medicine, Department of Molecular Genetics & Biochemistry, Pittsburgh, PA; 3) University of Pittsburgh, Graduate School of Public Health, Department of Environmental and Occupational Health, Pittsburgh, PA; 4) National Institute of Neuroscience, NCNP, Tokyo, Japan.

   Inherited biochemical deficits in patients with muscular dystrophy result in cycles of degeneration and regeneration of myofibers. Complementation of the biochemical deficiency in myofibers has been the major focus of experimental gene delivery in animal models of dystrophin and sarcoglycan deficiency. Recombinant adeno-associated virus (AAV) has been shown to deliver genes to mature normal muscle with long-term persistence, although persistence in dystrophic muscle has been more variable. We hypothesized that the degeneration/regeneration cycles and immune cell infiltrate in dystrophic muscle could be immunostimulatory, leading to increased immune response to a transgene relative to normal muscle. To test this hypothesis, we delivered both marker (b-galactosidase) and therapeutic genes (b-sarcoglycan) to dystrophic muscle and to normal muscle using identical AAV vectors. We show that dystrophic muscle does indeed elicit a stronger immune response to b-galactosidase, than normal muscle, showing both greater circulating antibodies (p< 0.05) and greater loss of transduced fibers (p< 0.05). Co-delivery of marker and therapeutic AAV attenuated the immune response (p<0.05), despite the use of twice as much AAV. Our results show that pathological muscle is immunostimulatory towards transgene-delivered proteins, leading to decreased persistence after AAV delivery. However, biochemical rescue of myofibers by a therapeutic transgene can attenuate this immune response, and in most cases leads to immunotolerance and long-term survival of transduced fibers.